{# load django_bootstrap5#%} CCDB server
Prof. Itay Mayrose Lab - Plant Evolution, bioinformatics, & comparative genomics


   CCDB is a community resource. If you are aware of any missing data or if any of the counts are erroneous please submit it to the forms submit new counts, or report erroneous counts.


CCDB → Angiosperms → Poaceae → Lolium → Lolium multiflorum Lam.

100 chromosome counts in Lolium multiflorum Lam.:


Name Accepted Name Gametophytic(n) Sporophytic(2n) Data Source reference
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Slovakia Database Májovský J. et al., 1976: Index of chromosome numbers of Slovakian flora (Part 5). - Acta Fac. Rerum Nat. Univ. Comen., Bot. 25: 1-18.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+1B   Flora Iberica Contribucin al conocimiento cariolgico de la subfam. Pooideae (Poaceae) en el SW de Espaa. Devesa, J. A. & T. Luque.Bol. Soc. Brot. ser. 261: 281-304(1988).
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Flora Iberica Contribution la connaissance cytotaxinomique des spermatophyta du Portugal. I. Gramineae Fernandes, A. & M. Queirs.Bol. Soc. Brot. ser. 243: 20-140(1969).
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   Flora Iberica Nmeros cromosmicos para la flora espaola, 321-330. Luque, T., C. Romero & J. A. Devesa.Lagascalia12(2): 286-289(1984).
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=4460
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Poland Database Skali?ska M., Pogan E., Czapik R. et al. 1978. Further studies in chromosome numbers of Polish Angiosperms. Twelfth contribution. Acta Biol. Cracov. Ser. Bot. 21, 31-63.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+1B   IPCN online Devesa, J. A. & T. Luque. 1988. Contribución al conocimiento cariológico de la subfam. Pooideae (Poaceae) en el SW de España. Bol. Soc. Brot., sér. 2 61: 281–304.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Buckner, R. C., P. B. Burrus & G. C. Eizenga. 1985. Genetic, morphological, and agronomic characteristics of Lolium-Festuca amphiploids. Crop Sci. (Madison) 25: 757–761.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN online Mehra, P. N. 1982. Cytology of East Indian grasses. ý.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Probatova, N. S. & A. P. Sokolovskaya. 1983. Chromosome numbers in Adoxaceae, Chloranthaceae, Cupressaceae, Juncaceae, Poaceae. Bot. Zhurn. SSSR 68 (12): 1683.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Schifino, M. T. & H. Winge. 1983. Karyotypes and nuclear DNA content of species of the Briza complex and some other genera of Poeae (Gramineae). Revista Brasil Gen;aaet. 6: 245–259.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN online Luque, T., C. R. Zarco & J. A. Devesa. 1984. Números 321-330. In Números cromosómicos para la flora española. 300-364. Lagascalia 12: 286–290.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+0-2B   IPCN online Spies, J. J. & S. Voges. 1988. Chromosome studies on African plants. 7. Bothalia 18: 114–119.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN online Devesa, J. A., T. Luque & P. Gómez. 1990. Números cromosomáticos de plantas occidentales, 591--601. Anales Jard. Bot. Madrid 47: 411–417.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Ghorai, A. & A. Sharma. 1981. Chromosome studies in some Festuceae. J. Indian Bot. Soc. 60: 148–153.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+1B   IPCN online Faruqi, S. A., H. B. Quraish & M. Inamuddin. 1987. Studies in Libyan grasses X. Chromosome number and some interesting features. Ann. Bot. (Rome) 45: 75–102.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Ko|3zuharov, S. I. & A. V. Petrova. 1991. Chromosome numbers of Bulgarian angiosperms. Fitologija 39: 72–77.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14, 28 IPCN online Wu, A. z. & D. x. Chen. 1991. The karyotype analysis of Oryzon ryegrass and its three lines. J. Shanghai Agric. Coll. 9(4): 253–259.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Cao, M. s., W. p. Chen & D. j. Liu. 1994. Cytogenetic studies of intergeneric hybrids F1 and amphiploid between Lolium multiflorum Lam. and Festuca arundinacea var. glaucescens Boiss. Sci. Agric. Sin. 27(5): 69–76.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Griffiths, P. D., H. J. Oughan & R. N. Jones. 1994. Genotypic and environmental effects on endopolyploidy in the epidermal tissues of Lolium perenne L. and Lolium multiflorum L. New Phytol. 128: 339–345.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN online Spies, J. J., S. M. C. v. Wyk, I. C. Nieman & E. J. L. Liebenberg. 1997. Cytogenetic studies in some representatives of the subfamily Pooideae (Poaceae) in South Africa. 3. The tribe Poeae. Bothalia 27: 75–82.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN online Spies, J. J., T. H. Burger & S. M. C. v. Wyk. 1999. Chromosome studies on African plants. 12. The tribes of subfamily Pooideae. Bothalia 29(2): 335–341.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Valdés, B., R. Parra, R. Parrilla & C. Reina. 1998. Números cromosómicos de plantas de Marruecos. Lagascalia 20(2): 223–230.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7 14 IPCN online Rees, H. & P. J. Dale. 1974. Chiasmata and variability in Lolium and Festuca populations. Chromosoma 47: 335–351.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Skalinska, M. 1978. Further studies in chromosome numbers of Polish angiosperms. Twelfth contribution. Acta Biol. Cracov., Ser. Bot. 21: 31–63.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7 14 IPCN online Evans, G. M. & A. J. Macefield. 1974. The effect of B chromosomes on homoeologous pairing in species hybrids. II. Lolium multiflorum X Lolium perenne. Chromosoma 45: 369–378.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Ghorai, A. & A. Sharma. 1981. Chromosome studies in some Festuceae. J. Indian Bot. Soc. 60: 148–153.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Jauhar, P. P. 1975. Chromosome relationships between Lolium and Festuca (Gramineae). Chromosoma 52: 103–121.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN online Parkash, O. 1979. Cytological investigations in some grasses of northeastern India (Festucoids, Ischaemum Linn., Eulalia Kunth and Saccharum spontaneum Linn.). Ph.D. Thesis, Chandigarh. 1979.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14, 28 IPCN online Kotseruba, V. V. 1997. Ideogram of the chromosome banding pattern of Lolium multiflorum Lam. Pages 131--132 in Agrobiotechnology of Plants and Animals. Abstracts of an International Conference. Kiev.
!   Lolium perenne var. italicum Parnell Lolium multiflorum Lam.   14 IPCN online Pohl, R. W. & G. Davidse. 1971. Chromosome numbers of Costa Rican grasses. Brittonia 23(3): 293–324.
!   Lolium perenne var. italicum Parnell Lolium multiflorum Lam.   14 IPCN online Pohl, R. W. & G. Davidse. 1971. Chromosome numbers of Costa Rican grasses. Brittonia 23(3): 293–324.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN66 Malik&Thomas 1966a, b
!   Lolium westerwoldicum Breakw. Lolium multiflorum Lam.   14 IPCN66 Malik&Thomas 1966b.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14+4B IPCN72 DELAY, J., & D. PETIT. 1972. Chromosomes surnumeraires chez des Graminees. Inf. Ann. Caryosyst. et Cytogenet. 6: 1-4.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN67-71 FERNANDES, A . & M. QUEIROS. 1969. Contribution a la connaissance cytotaxinomique des Spermatophyta du Portugal. I. Gramineae. Bol. Soc. Brat. 43(ser.2l: 3-140.
!   Lolium perenne var. italicum Parnell Lolium multiflorum Lam.   14 IPCN67-71 POHL, R.W., & G. DAVIDSE. 1971. Chromosome numbers of Costa Rican grasses. Brittonia 23: 293-324.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN75-78 Mehra & Sharma 1975b
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN75-78 Skalinska et al. 1978
!   Lolium multiflorum subsp. italicum (A. Braun) Schinz & R. Keller Lolium multiflorum Lam.   14 IPCN75-78 Ferakova 1976
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN75-78 Uhrikova 1978
!   Lolium multiflorum subsp. italicum (A. Braun) Schinz & R. Keller Lolium multiflorum Lam.   14 Love, 1948 Peto 1933.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   IPCN73-74 MEHRA, P.N., & P. REMANANDAN. 1973. Cytological investigations on W.l Himalayan Pooideae. Cytologia 38: 237-258.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 IPCN73-74 RAICU, P., & RODICA-CHIRILA. 1971. Caryotype et idiogramme de deux populations Roumaines de Lolium perenne L. et Lolium multiflorum Lamk Ann. Amelior. Pl. 21: 67-74.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Kumar, 1987 Evans, G. 1926 ; Tischler, G. 1934 ; Heiser, C.B. & Whitaker, T.W. 1948 ; Crowder, L.V. 1953 ; Hubbard, C.E. 1954 ; Tateoka, T. 1954 ; Shibata, K. 1956; Hovin, A.W. el a/. 1963 Hovin, A.W. & Hill, H.D. 1966 '; Malik, C .P. & Thomas, P.T. 1966, 1967; Mehra, P.N. et al. 1968 Fernandes, A. & Queiros, M. 1969 ; Mehra, P.N. & Sunder, S. 1969 ; Mehra, P.N. & Remanandan, P. 1973a ; Mehra, P.N. & Sharma, M.L. 1975c I4+4B Delay, J. & Petit, D. 1972
!   Lolium italicum A. Braun Lolium multiflorum Lam.   14 Kumar, 1987 Avdulov, N.P. (Fedorov) ; Jenkin, T.J. & Thomas, P.T. 1938 ; Tateoka,. T. 1955
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 eFlora  
!   Lolium italicum A. Braun Lolium multiflorum Lam.   14 C-values database Rees H, Jones GH. 1967. Chromosome evolution in Lolium. Heredity 22: 1-18.
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 C-values database Smarda P, Bures P, Horova L, Foggi B, Rossi G. 2008. Genome size and GC content evolution of Festuca: Ancestral expansion and subsequent reduction. Annals of Botany 101: 421-433.
!   Lolium italicum A. Braun Lolium multiflorum Lam.   14 Cave1956 Tateoka a
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Cave1956 Shiba.ta
!   Lolium multiflorum Lam. Lolium multiflorum Lam.   14 Cave1956supp Tateoka 1955
!   Lolium multiflorum Lam. Lolium multiflorum Lam. 7   Cave1963 Hovin et al. 1963
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+1B   CromoCat 2015 Devesa, J. A. & Luque, T. (1988). Contribución al conocimiento cariológico de la subfam. Pooideae (Poaceae) en el SW de España. Bol. Soc. Brot. 61: 281-304.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+1B   CromoCat 2015 Faruqi, S. A., Quraish, H. B. & Inamuddin, M. (1987). Studies in Libyan grasses X. Chromosome number and some interesting features Annali di Botanica 45: 75-102
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7+(0-2)B   CromoCat 2015 Spies, J. J. & S. P. Voges. (1988). Chromosome studies on African plants. 7. Bothalia 18: 114-119.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Devesa, J. A., Luque, T. & Gómez, P. (1990c). Números cromosomáticos de plantas occidentales, 591-601. Anales Jard. Bot. Madrid 47: 411-417.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Evans, G. M. & Macefield, A. J. (1974). The effect of B chromosomes on homoeologous pairing in species hybrids. II. Lolium multiflorum X Lolium perenne. Chromosoma 45: 369-378.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Luque, T., Romero Zarco, C. & Devesa Alcaraz, J. A. (1984). Números cromosómicos para la flora española, 321-330. Lagascalia 12(2): 286-289.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Mehra, P. N. (1982). Cytology of East Indian grasses
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Rees, H. & P. J. Dale. (1974). Chiasmata and variability in Lolium and Festuca populations. Chromosoma 47: 335-351.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Spies, J. J. & H. Du Plessis. (1986). Chromosome studies on African plants. 1. Bothalia 16: 87-91.
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Spies, J. J., T. H. Burger & S. M. C. van Wyk (1999). Chromosome studies on African plants. 12. The tribes of subfamily Pooideae Bothalia 29(2): 335-341
  Lolium multiflorum Lam. Lolium multiflorum Lam. 7   CromoCat 2015 Spies, J.J., S.M.C. Van Wyk, I.C. Nieman & E.J.L. Liebenberg (1997). Cytogenetic studies in some representatives of the subfamily Pooideae (Poaceae) in South Africa. 3. The Tribe Poeae Bothalia 27:75-82
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Avdulov, N. P. (1928). Systematicheskaya kariologiya semeestva Gramineae. Drievnik vsesojuznogo Sezda Bot. Leningrade 1928: 65-66.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Avdulov, N. P. (1931). Karyo-systematische Untersuchungen der Familie Gramineen. Bull. Appl. Bot., Genet. & Plant Breeding Suppl. 43: 1-438.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Buckner, R.C., Burrus, P.B. & Eizenga, G.C. (1985). Genetic, morphological, and agronomic characteristics of Lolium-Festuca amphiploids. Crop Science (Madison) 25: 757-761
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Cao, M.-s., Chen, W.-p. & Liu, D.-j. (1994). Cytogenetic studies of intergeneric hybrids F1 and amphiploid between Lolium multiflorum Lam. and Festuca arundinacea var. glaucescens Boiss. Scientia Agricultura Sinica 27(5): 69-76
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Cooper, J. P. & Saeed, S. W. (1949). Studies on growth and development in Lolium. I. Relation of the annual habit to head production under various systems of cutting. J. Ecol. 37: 233-259.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Crowder, L. V. (1953). Interspecific and intergeneric hybrids of Festuca and Lolium. J. Heredity 44: 195-204.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Evans, G. (1927). Chromosome complements in grasses. Nature 118: 841.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Evans, G. M. & Macefield, A. J. (1974). The effect of B chromosomes on homoeologous pairing in species hybrids. II. Lolium multiflorum X Lolium perenne. Chromosoma 45: 369-378.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Fernandes, A. & Queirós, M. (1969). Contribution à la connaissance cytotaxinomique des spermatophyta du Portugal. I. Gramineae. Bol. Soc. Brot. 43: 20-140.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Ghorai, A. & Sharma, A. (1981). Chromosome studies in some Festuceae Journal of the Indian Botanical Society 60: 148-153
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Griffiths, P. D., Oughan, H. J. & Jones, R. N. (1994). Genotypic and environmental effects on endopolyploidy in the epidermal tissues of Lolium perenne L. and Lolium multiflorum L. New Phytologist 128: 339-345
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Hovin, A. W. & Hill, H. D. (1966). B-chromosomes, their origin and relation to meiosis in interspecific Lolium hybrids. Amer. J. Bot. 53: 702-708.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Jauhar, P. P. (1975). Genetic regulation of diploid-like chromosome pairing in the hexaploid species Festuca arundinacea Schreb. and F. rubra L. (Gramineae). Chromosoma 52: 363-382.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Jenkin, T. J. & Thomas, P. T. (1938). The breeding affinities and cytology of Lolium species. J. Bot. 76: 10-12.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Jenkin, T. J. (1931). The method and technique of selection breeding and strain building in grasses. Bull. Bur. Pl. Genet. 3: 5-34.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Kozuharov, S. & Petrova, A. (1991). Chromosome numbers of Bulgarian Angiosperms. Fitologija 39: 72-77.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 László, H. & Marianna, V. K. (1971). A Lolium-faok diploid és tetraploid alakjainak összehasonlí tó vizsgálata. II. Az olasz perje (Lolium multiflorum Lam.). Növénytermelés 20: 205-212.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Nielsen, E. L. & L. M. Humphrey. (1937). Grass studies. I. Chromosome numbers in certain members of the tribes Festucae, Hordeae, Aveneae, Agrostideae, Chlorideae, Phalarideae and Tripsaceae. Amer. J. Bot. 24: 276-279.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Parkash, O. (1979). Cytological investigations in some grasses of northeastern India (Festucoids, Ischaemum Linn., Eulalia Kunth and Saccharum spontaneum Linn.). Ph.D. Thesis, Chandigarh. 1979.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Peto, F. H. (1933). The cytology of certain intergeneric hybrids between Festuca and Lolium. J. Genet. 28: 113-156.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Probatova, N. S. & A. P. Sokolovskaya. (1983). New chromosome numbers for vascular plants from the Islands of Peter the Great Bay (Primorye Territory). J. Bot. URSS 68(12): 1655-1662.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Probatova, N. S. & A. P. Sokolovskaya. (1984b). Chromosome numbers of the representatives of the families Butomaceae, Papaveraceae, Poaceae from the U.S.S.R. far East. U.S.S.R. (Komiassr and the Tumen' district). Bot. Žurn. 69: 410-412.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Rees, H. & P. J. Dale. (1974). Chiasmata and variability in Lolium and Festuca populations. Chromosoma 47: 335-351.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Schifino, M.T. & H. Winge. (1983). Karyotypes and nuclear DNA content of species of the Briza complex and some other genera of Poeae (Gramineae). Revista Brasil Gen;aaet. 6: 245-259.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Shalygin, J. N. (1941). Production of tetraploids in Lolium by treating germinating seeds with colchicine. Dokl. Akad. Nauk. SSSR. 30: 527-529.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Shibata, K. (1956). Karyotype analysis in some forage grasses. Bot. Mag., Tokyo 69: 165-168.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Shibata, K. (1957a). Karyotype analysis in some forage grasses. II. Jap. J. Genet. 32: 259-260.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Skalinska, M., E. Pogan, R. Czapik. (1978). Further studies in chromosome numbers of Polish Angiosperms. XII. Acta Biol. Cracov., Ser. Bot. 21: 31-63.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Tateoka, T. (1955a). Karyotaxonomic studies in Poaceae. II. Natl. Inst. Genet. (Japan), Ann. Rep. 5: 68-69.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Tateoka, T. (1955b). Karyotaxonomy in Poaceae. III. Further studies of somatic chromosomes. Cytologia 20: 296-306.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Tateoka, T. (1956a). Karyotaxonomic studies in Poaceae. III. Natl. Inst. Genet. (Japan), Ann. Rep. 6: 73-74.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Tateoka, T. (1956b). Karyotaxonomic in Poaceae. IV. Chromosomes and systematic relationship of several species. Bot. Mag., Tokyo 69: 112-117.
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Tateoka, T. (1959). Introduction to grasses. Tokyo
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Valdés, B., R. Parra, R. Parrilla & C. Reina (1998). Números cromosómicos de plantas de Marruecos Lagascalia 20(2): 223-230
  Lolium multiflorum Lam. Lolium multiflorum Lam.   14 CromoCat 2015 Wu, A. -Z. & D. -X. Chen (1991). The karyotype analysis of Oryzon ryegrass and its three lines. J. Shanghai Agric. Coll. 9(4):253-259
  Lolium multiflorum Lam. Lolium multiflorum Lam.   28 CromoCat 2015 Wu, A. -Z. & D. -X. Chen (1991). The karyotype analysis of Oryzon ryegrass and its three lines. J. Shanghai Agric. Coll. 9(4):253-259